A stapling system is disclosed. The stapling system comprises a housing comprising a rotary drive member, an elongate shaft extending from the housing, wherein the elongate shaft defines a longitudinal axis, and an end effector comprising a jaw configured to support a staple cartridge and an anvil rotatable relative to the jaw between an open position and a fully-closed position, wherein the anvil comprises a cam surface. The stapling system further comprises a closure cam operably coupled with the rotary drive member, wherein the closure cam is configured to move longitudinally to engage the cam surface and transmit a closing motion to the anvil to move the the into the fully-closed position and an opening member configured to move longitudinally to apply an opening force to the anvil at a location other than the cam surface to move the anvil into the open position.
