The present invention provides wheat grain of the species Triticumaestivum, the grain comprising i) mutations in each of its SSIIa genes such that the grain is homozygous for a mutation in its SSIIa-A gene, homozygous for a mutation in its SSIIa-B gene and homozygous for a mutation in its SSIIa-D gene, wherein at least two of the mutations in said SSIIa genes are null mutations, mutations in each of its SSIIIa genes such that the grain is homozygous for a mutation in its SSIIIa-A gene, homozygous for a mutation in its SSIIIa-B gene and homozygous for a mutation in its SSIIIa-D gene, wherein at least two of the mutations in said SSIIIa genes are null mutations ii) a total starch content comprising an amylose content and an amylopectin content, iii) a fructan content which is increased relative to wild-type wheat grain on a weight basis, preferably between 3% and 12% of the grain weight, iv) a β-glucan content, v) an arabinoxylan content, vi) a cellulose content. The grain has a weight of between 25mg and 60mg, and the amylose content is between 45% and 70% on a weight basis of the total starch content of the grain as determined by iodine binding assay. The amylopectin content on a weight basis is reduced relative to the wild-type wheat grain, and each of the β-glucan content, arabinoxylan content and cellulose content are increased relative to the wild-type wheat grain on a weight basis, such that the sum of the fructan content, β-glucan content, arabinoxylan content and cellulose content is between 15% and 30% of the grain weight.La présente invention concerne un grain de blé de l'espèce Triticumaestivum, le grain comprenant i) des mutations dans chacun de ses gènes SSIIa de telle sorte que le grain est homozygote pour une mutation dans son gène SSIIa-A, homozygote pour une mutation dans son gène SSIIa-B et homozygote pour une mutation dans son gène SSIIa-D, au moins deux des mutations dans lesdits gènes SSIIa étant des mutations nulles, des mutations dans chacun de
