A fluid dispenser head for mounting on a fluid reservoir so as to constitute a dispenser. The head includes a fluid dispenser member with a body and a valve rod. A dispenser endpiece is rotatably mounted on the valve rod and a turning pusher is manually axially movable so as to move the dispenser endpiece and the valve rod. A transmission part is interposed between the endpiece and the pusher and secured to the pusher in axial movement. An actuator mechanism causes the endpiece and pusher to turn, but not the transmission part. The actuator mechanism moves the transmission part axially between a non-working, storage position and a working, actuation position, in such a manner as to cause the pusher to move axially. The part transmits directly and axially to the endpiece, any thrust force exerted on the pusher in the working position.