The present invention relates to nucleic acids encoding flavonoid biosynthetic enzymes, more particularly flavonoid biosynthesis-regulating transcription factors, and the use thereof for the modification of flavonoid biosynthesis in plants. The present invention also relates to constructs and vectors including such nucleic acids, and related polypeptides. More particularly, the present invention relates to a PURPLE ANTHOCYANIN PIGMENT 1 (PAP1) or PAP1-like transcription factor.