The present invention relates to non-transgenic and transgenic plants, preferably crop plants, comprising at least one mutation causing a substitution of an amino acid within the CATD domain of the centromere histone H3 (CENH3), preferably within the α2-helix of the CATD domain, which have the biological activity of a haploid inducer. Further, the present invention provides methods of generating the plants of the present invention and haploid and double haploid plants obtainable by crossing the plants of the present invention with wildtype plants as well as methods of facilitating cytoplasm exchange.
