A surgical instrument assembly is disclosed which comprises a shaft, an articulation joint, and an end effector rotatably connected to the shaft about the articulation joint. The end effector comprises a first jaw, a second jaw movable relative to the first jaw, a staple cartridge comprising a plurality of staples removably stored therein, and an anvil configured to deform the staples. The end effector further comprises a drive input, a closure system configured to move the second jaw between an open position and a closed position, a staple firing system configured to eject the staples from the staple cartridge, and a transmission positioned in the end effector, wherein the transmission is configurable in three operating states for operably coupling the closure system and the staple firing system with the drive shaft.