The invention provides recombinant DNA molecules and constructs, and their nucleotide sequences, useful for modulating gene expression in plants. The invention also provides transgenic plants, plant cells, plant parts, and seeds comprising a recombinant DNA molecule comprising a DNA molecule operably linked to a heterologous transcribable DNA molecule, as well as methods of their use.La invencion proporciona las moleculas y constructos de ADN recombinante y sus secuencias de nucleotidos, utiles para modular la expresion genica en las plantas； la invención tambien proporciona plantas transgenicas, celulas de planta, partes de planta, y semillas que comprende una molecula de ADN recombinante que comprende una molecula de ADN unida operativamente a una molecula de ADN transcribible heterologa, asi como metodos de su uso.
