A surgical instrument with separate closure and firing systems for applying closure and firing motions. In one form, the surgical instrument includes a hollow closure member and a firing member. The hollow closure member is configured to move axially between an unactuated position and closure positions. The firing member is supported for axial travel between a starting position in the hollow closure member and an ending position in the surgical end effector. The closure system is configured to apply closure actuation motions to the hollow closure member. The firing system is configured to apply a firing motion to the firing member in order to move the firing member from the starting position to the ending position.
