An airflow directional control system (100) in an agricultural harvester (10) having a grain tank (28) or other structure, an internal combustion engine (32) and a cooling system (70) cooler box (71) disposed between the grain tank (28) or other structure and the internal combustion engine (32) includes a cooler box cover (102, 110) over the cooler box (71) between the grain tank (28) or other structure and the internal combustion engine (32), at least one air director (120) between the cooling box (71) and the internal combustion engine (32) and a panel adjustment system (130) controlling tilt of side panels (132, 134) of the harvester (10).