The present invention relates to a dispenser (200) for dispensing a dose of a gaseous, gas borne or droplet substance from a substance source, the dispenser comprising: a body for receiving a substance source, the body having a mouthpiece; and a dose counter (203) for indicating a number of actuations of the dispenser corresponding to a number of doses dispensed from a substance source, or a number of doses remaining in a substance source, the dose counter comprising: a counter having indicia, the counter being rotatable within the body in increments about the longitudinal axis of the body; a counter driver (206) for driving the counter, the counter driver being arranged to be reciprocatably moveable within the body in the longitudinal axis in response to an actuation of the dispenser; and a drive mechanism (205) for rotating the counter, the drive mechanism being coupled to the counter driver and configured to rotate the counter in response to longitudinal movement of the counter driver, wherein the body comprises a counter driver guide (230) configured to guide the counter driver (206) in the body so as to prevent rotation of the counter driver in the longitudinal axis. Examplary means to prevent rotation is the counter guide (230) interacting with notched portion (220) on counter driver.