In various embodiments, a shaftless end effector is disclosed. The end effector can comprise an anvil assembly including a staple forming surface. The end effector can also comprise a staple cartridge channel configured to receive a staple cartridge therein. The end effector can include a closure system configured to close the anvil. In various instances, the closure system can also form the staples contained within the staple cartridge. The end effector can also include a cutting member configured to cut tissue positioned intermediate the anvil and the staple cartridge. In various instances, the cutting member can be actuated by the closure system. In other instances, the end effector can include an independent system for actuating the cutting member. In various instances, the end effector can be delivered to a surgical site through a trocar and then operated by one or more actuators that are assembled to the end effector.