Provided are isolated polynucleotides and polypeptides. Also provided are recombinant DNA constructs useful for conferring improved tolerance in plants to insect pests, compositions (such as plants or seeds) comprising these recombinant DNA constructs, and methods utilizing these recombinant DNA constructs. The recombinant DNA constructs comprise a polynucleotide operably linked to a promoter that is functional in a plant, wherein said polynucleotides encode insect tolerance polypeptides.