A surgical instrument including a shaft and an end effector. The end effector may comprise a first jaw and a second jaw. The first jaw may be movable relative to the second jaw between an open configuration and a closed configuration. The surgical instrument may comprise a closure assembly which may be operably engaged with the first jaw. The closure assembly may comprise a closure trigger and a dampening system. The closure trigger may be configured to be actuated from a first position to a second position to close the first jaw. The dampening system may be configured to retard the opening of the closure trigger. The dampening system may comprise an aperture configured to receive the projection at a first end of the aperture and a seal configured to form a fluid seal between said projection and said aperture sidewall.